PER CURIAM.

We have on appeal the judgment and sentence of the trial court imposing the death penalty upon Gary Lee Thorp. We have jurisdiction. Art. V, § 3(b)(1), Fla. Const. For reasons which follow, we reverse appellant's conviction for first-degree murder because we find harmful error in the admission of improper evidence at trial and remand for a new trial.

MATERIAL FACTS

Gary Thorp was convicted of first-degree murder and sentenced to death for the murder of Sharon Chase. The record reflects that Chase's nude body was discovered in the bushes in Bean Park on June 24, 1993. Her body was lying face up, with her legs spread apart and knees partially bent. Her shirt had been pulled down to her waist, and her shorts were discovered some thirty feet away from the body. Semen discovered inside the victim's vagina matched Thorp's DNA profile. The medical examiner opined that death was caused by strangulation. Thorp subsequently was arrested for the murder of Sharon Chase in October of 1996.

Evidence presented at trial indicates that at approximately midnight on June 23, 1993, the victim and a white male were seen together crossing a bridge into Bean Park near U.S. 1 in Melbourne, Florida. The witness, Paul Symeon, was fishing in the marina across the street from the park and observed Chase and a man enter the park. Symeon described the man as "skinny or lanky," taller than the victim, with a dark complexion and dark, wavy hair. Symeon testified that the two "milled around for a while" and were then approached by another unidentified couple. After approximately ten minutes, Symeon heard rustling sounds in the bushes for a couple of minutes and then heard a sound similar to a moan. When he turned to look, he could not see what was causing the noises. However, Symeon noticed that no one was in the park at that time. At trial, Symeon identified Chase as the woman he had seen in the park. However, Thorp did not fit the physical description given by Symeon of the man he saw with Chase and Symeon could not identify Thorp as being present in the park the night of the crime.

Other evidence at trial revealed that Thorp and another man, William Deering, had been living at the Christ Is The Answer (CITA) Mission in Melbourne. On the night of the homicide, Thorp and Deering both missed the 11 p.m. bed check at the mission and, as a result, were not permitted to sleep there that night. David Gallamore, an employee at the mission, saw Thorp around 1:15 a.m. and noticed Thorp had been drinking and had blood on his shirt and bruises on his knuckles. Gallamore asked Thorp why he missed the earlier bed check. Thorp stated that he had been in a fight at a Burger King restaurant. He also asked Gallamore if Deering had been at the mission that night. In fact, Gallamore saw Deering at the mission at around 12:30 a.m.

Finally, the State presented Timothy Bullock, an inmate housed with Thorp at the Brevard County Jail during the spring of 1994 where Thorp was serving time for an unrelated crime. Bullock testified that Thorp told him that he and another man "took a hooker down by the bridge and did her," during which he got a lot of blood on himself. Bullock testified that Thorp admitted that he expected to be blamed for the Chase murder. In addition, and over defense counsel's objection, Bullock was allowed to testify that he interpreted Thorp's statement that he "did a hooker" to mean that Thorp killed her. Finally, Bullock testified that Thorp had admitted that, in order to be allowed into the mission with blood on his clothes that night, he had told the mission employee that the blood was caused by a fight at Burger King.

The defense did not present any witnesses during the guilt phase of the trial. After deliberation, the jury found Thorp guilty of first-degree murder. During the penalty phase, the State presented two witnesses, the prosecutor and the lead investigating officer in a 1994 murder case in which Thorp pled nolo contendere to a charge of second-degree murder for the fatal stabbing of Randy Appleman. The defense presented several witnesses of their own who testified that Thorp got along with others, was a hard worker, and assisted the mission director as his "right hand man" at the mission where he lived. Both of Thorp's parents testified that Thorp was born prematurely, suffered from cerebral palsy as a child, and ultimately developed a serious drinking problem. Thorp testified on his own behalf about his difficult early life, his cerebral palsy, his struggles with drugs and alcohol, and his checkered educational and employment history. During his testimony Thorp admitted to having consensual sexual intercourse with the victim but denied killing her. Thorp also denied killing Appleman, although he admitted to stealing Appleman's property and forging some of Appleman's checks. Thorp expressed remorse for his past lifestyle but denied killing anyone. The jury recommended a sentence of death by a vote of ten to two, and the trial court followed this recommendation.

APPEAL

On appeal, Thorp raises seven issues for this court's review. We agree with Thorp that reversal is required on two of his claims: claim (1), concerning the trial court's failure to suppress evidence obtained from a substantially misleading affidavit for search warrant, and claim (3), concerning the trial court's error in allowing improper opinion testimony to go to the jury. Because we are reversing Thorp's conviction and sentence based on those asserted errors, we do not address claims (4), (5), (6), and (7) as they are now moot.

Sufficiency of the Evidence

Thorp argues the trial court erred in denying his motion for judgment of acquittal because the State's evidence failed to exclude the reasonable hypothesis of innocence that someone other than Thorp killed the victim. He contends that the DNA evidence proved only that he had sexual intercourse with the victim, not that he killed her. In addition, Thorp further contends that the evidence at trial contradicts the State's theory of how the murder occurred. Although we find prejudicial error with regard to two aspects of the State's case, we nevertheless find that the evidence was sufficient to overcome Thorp's motion for judgment of acquittal.

Because there were no eyewitnesses or other direct evidence of Thorp's commission of the murder, the State's case against Thorp was predicated chiefly upon circumstantial evidence. As we stated in State v. Law, 559 So.2d 187 (Fla. 1989):

A special standard of review of the sufficiency of the evidence applies where a conviction is wholly based on circumstantial evidence. Jaramillo v. State, 417 So.2d 257 (Fla. 1982). Where the only proof of guilt is circumstantial, no matter how strongly the evidence may suggest guilt, a conviction cannot be sustained unless the evidence is inconsistent with any reasonable hypothesis of innocence. McArthur v. State, 351 So.2d 972 (Fla. 1977); Mayo v. State, 71 So.2d 899 (Fla. 1954). The question of whether the evidence fails to exclude all reasonable hypotheses of innocence is for the jury to determine, and where there is substantial, competent evidence to support the jury verdict, we will not reverse. Heiney v. State, 447 So.2d 210 (Fla.), cert. denied, 469 U.S. 920, 105 S.Ct. 303, 83 L.Ed.2d 237 (1984); Rose v. State, 425 So.2d 521 (Fla. 1982), cert. denied, 461 U.S. 909, 103 S.Ct. 1883, 76 L.Ed.2d 812 (1983), disapproved on other grounds, Williams v. State, 488 So.2d 62 (Fla. 1986).

, 559 So.2d at 188 . On the basis of the above rule, we held in Law that where the State fails to present evidence from which the jury can exclude every reasonable hypothesis except that of guilt, the court should grant a motion for judgment of acquittal. Id.

Thorp contends that the State's evidence fails to exclude the possibility that someone other than Thorp killed the victim since several people were seen in the park on the night of the murder and none of the witnesses could actually identify Thorp as the person they observed walking with the victim prior to the murder. He also points out there is no physical evidence that Thorp killed the victim.

The State's principle evidence linking Thorp to the crime includes the DNA evidence, Thorp's statements to his cellmate Bullock, and Thorp's physical appearance and condition on the night of the crime (i.e., that he was observed with blood on his clothing). We conclude that this evidence was sufficient to require this case to be submitted to the jury.

As noted above, the DNA evidence indicates that Thorp was with the victim and had sexual intercourse with her the night of the murder. We recognize that while the DNA evidence, like fingerprint evidence, does not conclusively prove that Thorp committed the murder, cf. Jaramillo v. State, 417 So.2d 257, 257 (Fla. 1982) (disregarding fingerprint evidence where State failed to prove the defendant's fingerprints could only have been placed on items in victim's home at time murder was committed), the DNA evidence supports the State's contention that Thorp was with the victim in the park around the time she was killed.

The other significant evidence against Thorp is his supposed confession of the murder to cellmate Timothy Bullock. While Bullock testified that Thorp admitted he expected to be blamed for the murder of a prostitute, Bullock did not testify that Thorp ever directly admitted to killing the victim. His actual statement, according to Bullock was that Thorp and another man "did a hooker." However, direct evidence was also presented that Thorp was seen with injuries and blood on his clothes on the night of the crime, injuries that could be consistent with a physical struggle with the murder victim who had considerable bruises and abrasions on her body even if she did not bleed extensively. Obviously, the victim's assailant could also have been injured and bled.

On the face of the record before us, we find this evidence legally sufficient to convict Thorp of first-degree murder. While the State's evidence may not have been conclusive, it strongly suggests that Thorp struggled with and killed the victim. It was up to the jury to determine if this evidence proved Thorp's guilt beyond a reasonable doubt. Accordingly, we find no error in the trial court denying Thorp's motion for judgment of acquittal.

Motion to Suppress

On the other hand, we find the trial court committed prejudicial error by failing to exclude Thorp's blood samples and the resulting DNA evidence obtained from those samples. The record in this case reflects that approximately a year after the murder occurred, the police obtained a search warrant to take a blood sample from Thorp. The affidavit filed by the police stated that Thorp matched the description of the person Symeon observed walking with the victim on the night of the murder. The affidavit further provided the following facts: that Chase was a known prostitute who worked in the Melbourne area; that on the night of the murder Symeon heard moaning (muffled screams) emanating from the park after the suspect and victim had entered it; that semen was discovered in the vaginal cavity of the victim in an amount sufficient to conduct a DNA comparison; that Thorp was the primary suspect in a unrelated stabbing incident in which the victim was also found nude; that Thorp frequented the Melbourne area near Bean Park and had lived at the CITA mission; that Thorp despised prostitutes; that Thorp and Deering had not spent the night at the mission on the night of the murder; that one of the them was observed with blood on his clothes; and that the suspect was observed wearing a ball cap and that Thorp consistently wore ball caps. Based on these facts, the magistrate judge issued a search warrant to obtain two vials of Thorp's blood for the purpose of conducting a DNA comparison with the semen recovered from the victim.

During pretrial proceedings, Thorp moved to suppress the physical evidence obtained as a result of the search warrant. At an evidentiary hearing on the matter, Thorp asserted the affidavit contained false statements and material omissions which, when considered in light of the remaining factual allegations, defeated a finding of probable cause. The State hesitantly conceded that the statement concluding that the suspect matched Thorp's description and the statements concerning the existence of a ball cap should be redacted. It maintained, however, that the remaining factual allegations in the affidavit were sufficient to establish probable cause. The trial court found that the affidavit falsely indicated that the description given by Symeon matched Deering and Thorp. However, the court concluded that the misstatement was not willful and, even if it was removed, the remaining content of the affidavit established probable cause. We disagree with the trial court's conclusion.

If an affidavit for search warrant contains intentional false statements or statements made with reckless disregard for the truth, the trial court must excise the false material and consider whether the affidavit's remaining content is sufficient to establish probable cause.See Franks v. Delaware, 438 U.S. 154, 156 (1978); Terry v. State, 668 So.2d 954, 958 (Fla. 1996). This rule contains two components. First, the trial court must determine whether the affidavit contains an intentional false statement or a statement made in reckless disregard for the truth. Mere neglect or statements made by innocent mistake are insufficient. See Franks, 438 U.S. at 171. Second, if the court finds the police acted deceptively, the court must excise the erroneous material and determine whether the remaining allegations in the affidavit support probable cause. See id. at 171-72. If the remaining statements are sufficient to establish probable cause, the false statement will not invalidate the resulting search warrant. See Terry, 668 So.2d at 958. If, however, the false statement is necessary to establish probable cause, the search warrant must be voided and the evidence seized as a result of the search excluded. See id. (citing Franks, 438 U.S. at 156).

Thorp points to several critical and substantial misstatements set out in the affidavit, the falsity of which were known to the police officer affiant. The first pertains to the identification of Thorp as the person last seen with the victim. The affidavit states that "the description of the male that was seen walking with the female (that is most probably Sharon Chase), prior to the murder, is consistent with the description of Thorpe and Deering." As noted above, the State conceded that the statement in the affidavit that Thorp and Deering matched the description given by Symeon should be stricken from the affidavit. The affidavit describes Thorp as a white male, approximately six feet tall, 220 pounds, with blue eyes and bald/brown hair. Deering is a white male, approximately five feet, eight inches tall, 165 pounds, with brown eyes and brown hair. According to the statement Paul Symeon gave to the police shortly after the victim's body had been discovered, however, the person he observed with the victim was a white male, approximately six feet tall, seemingly thin, with medium to long dark brown hair, and adark complexion. Symeon initially described the suspect as a minority and later, during a hypnotic session with the police, characterized the suspect as possibly Spanish. None of these known and obviously relevant facts about the suspect's complexion were included in the affidavit. Moreover, the officer who prepared the affidavit improperly likened Thorp to the person last seen with the victim. From the descriptions above, there appears to be little or no similarity between Thorp, Deering, and the person Symeon described as walking with the victim prior to the murder. Based on the patent differences in physical characteristics of the three persons involved and the crucial importance of this information in establishing probable cause, we find at least a patent and reckless disregard for the truth in this misstatement. See Franks.

In addition to the above, we find the affidavit patently misstates Symeon's description of the events in the park that night. The affidavit states that Symeon heard "moaning (muffled screams)" emanating from the park. The parenthetical was added not by the witness Symeon, but by the police, and obviously was intended to give meaning to the word "moaning." However, Symeon never described the sounds he heard as "muffled screams." In fact, in his initial statement to the police, Symeon stated that the sound he heard was "definitely not a scream and it was not a moan, it was something in between that." While under hypnosis, Symeon again described the noise he heard as a "moan" but made no reference to "muffled screams." He never saw what caused the moans. Thus, the affiant improperly inserted a misstatement as to what Symeon heard, and the affidavit's reference to "muffled screams" should also have been redacted.

Before considering the affidavit absent the misstated facts, we must also address other relevant facts known to the police, but which Thorp claims were improperly omitted from the affidavit. Thorp's contention is that had the omitted facts been included in the affidavit for search warrant, they would have defeated a finding of probable cause. Omissions made with intent to deceive or with reckless disregard must also be considered in considering a motion to suppress. See Johnson v. State, 660 So.2d 648, 656 (Fla. 1995). The only difference between omitted facts and misstated facts is that in the case of omitted facts, the reviewing court must first determine whether the omitted facts, if added to the affidavit, would have defeated probable cause. See id. The reviewing court must further find that the omission resulted from intentional or reckless police conduct that amounts to deception. See id. "When a material fact is omitted from the affidavit filed in support of the probable cause determination, such fact constitutes a material omission if a substantial possibility exists that the omission would have altered a reasonable magistrate's probable cause determination." State v. Van Pieterson, 550 So.2d 1162, 1164 (Fla. 1st DCA 1989).

Here, several important and relevant facts were omitted which, when considered together with the other errors, appear to indicate a lack of probable cause. For example, the medical examiner had examined the victim's body shortly after it was discovered and concluded that the victim had not bled profusely. Yet this fact was not included in the affidavit. This omitted material is important in light of the fact the affidavit states that Thorp was seen with blood on his clothes, giving the impression that he had been involved in a bloody struggle. Had the magistrate been told the victim did not shed blood, the evidence that Thorp had blood on his clothes might have carried less weight in the probable cause determination.

Other omissions concern Symeon and the actual description he provided to the police. For example, the affidavit does not reveal that Symeon was hypnotized and that several of his statements relied upon in the affidavit were the result of a hypnotic session. One such statement induced during hypnosis concerns the fact that Symeon purportedly stated that the suspect was wearing a baseball cap. It is undisputed that Symeon never mentioned a baseball cap during his initial interview with the police. While under hypnosis, however, Symeon claimed that the man he saw with the victim was wearing a baseball cap. During that same session, he later reversed himself and stated that the suspect was not wearing a baseball cap or hat. The affidavit does not mention this patent inconsistency in Symeon's various statements to the police or the circumstances under which they were provided. These omissions are material because the affidavit also claims that Thorp consistently wears baseball caps, a fact obviously intended to strengthen the connection to Thorp as the perpetrator, but which becomes irrelevant if the perpetrator was not wearing a cap on the night of the murder.

We find that once the misstated facts are redacted and the omitted facts are included, the remaining factual allegations in the amended affidavit do not establish probable cause to believe that Thorp committed murder. Notably, despite the lengthy factual recitations in the affidavit, none of the truthful allegations identified Thorp as the person last seen with the victim or even placed Thorp in the park on the night of the homicide. At most, the affidavit indicates Thorp had been in an altercation that night and that he lived at the CITA mission. There is nothing within the affidavit's factual recitations to lead a magistrate to reasonably believe that Thorp was the person observed with the victim and was the person who caused the victim's death. Indeed, the affidavit also reveals that at least two other men were in the park that night, neither of whom was identified. On the strength of the amended affidavit, we cannot say the factual allegations therein are sufficient to establish probable cause to believe Thorp was involved in the instant murder. Accordingly, we conclude the trial court abused its discretion in denying Thorp's motion to suppress the blood samples and the DNA test results from those samples. See Jones v. State, 343 So.2d 921, 922 (Fla. 3d DCA 1977).

We must next determine whether the erroneous admission of the blood samples and resulting DNA analysis from those samples constitutes harmful error. See State v. DiGuilio, 491 So.2d 1129 (Fla. 1986). Upon the record in this case, we find that it does. The State presented three experts in DNA analysis. The first expert conducted a Restriction Fragment Length Polymorphisms (RFLP) test on the semen found in the victim's vagina and compared the results to Thorp's and Deering's blood samples. After comparing the known samples, the expert was able to exclude Deering as the semen donor but indicated a possible match with Thorp's DNA profile. Dan Nippes, the second expert, performed a different DNA analysis on the samples, known as the Polymerase Chain Reaction (PCR) test, and concluded that the semen found in the victim matched Thorp's DNA profile. Nippes further opined that the chance of such a match occurring in the population is approximately one in one thousand Caucasians. The third expert testified that the statistical probability of finding a male unrelated to Thorp who matched both the RFLP DNA test result and the PCR DNA test result would be approximately one in 3.6 billion. During closing arguments, the State relied on this evidence in arguing that there is no reasonable doubt that the semen found in the victim belonged to Thorp.

The only other significant evidence of guilt included Thorp's statements to a fellow inmate and a witness's testimony that Thorp had blood on this clothes on the night of the murder. Based on the conclusive nature of the DNA evidence, however, and because it is the only physical evidence placing Thorp at the scene of the crime, we cannot say beyond a reasonable doubt that the improperly admitted DNA evidence had no effect on the verdict in this case. See DiGuilio, 491 So.2d at 1136; see also Goodwin v. State, 751 So.2d 537, 546 (Fla. 1999) (reaffirming harmless error analysis in DiGuilio). Thus, the court's error in failing to exclude the blood samples and DNA test results was harmful error, requiring reversal.

Admission of Opinion Testimony

We further find harmful error in the admission of improper opinion testimony by a lay witness. Thorp contends the trial court erred in permitting Bullock to testify as to what he thought Thorp meant when Thorp told him that he (Thorp) "did a hooker." The State, on the other hand, argues that Thorp failed to preserve this claim for appellate review and, even if preserved, the claim is meritless. We disagree with both of the State's contentions.

At trial, the following colloquy occurred between the State and Bullock:

[State] Q. Did you ever have any conversations with Gary Lee Thorp about the death of the prostitute under the bridge?

[Bullock] A. Yes, I did.

Q. Did he indicate to you that he knew about this incident?

A. Yes, he did.

Q. And did he indicate to you that he expected that he might be charged with this?

A. He told me he expected to be.

Q. What did he expect the charge to be?

A. Murder.

Q. What did he say about that incident?

A. He told me that he knew the girl, that he had taken her down by the bridge and did her, that she was a hooker.

Q. Those were his words, that he did her?

A. Yes.

Q. What did you take that to mean?

A. That he had-

[Defense counsel]: I object. That's not relevant. That's speculation. This gentlemen has not been qualified to interpret anything that he thought was said. It's just irrelevant and immaterial.

[The court]: Overruled.

Q. You may answer the question. What did you take that to mean?

A. I took that to mean that he was the one that killed her.

Q. Did he indicate to you whether or not there was any other person associated with this incident?

A. Yes, he did.

Q. How did he describe that to you?

A. I believe his words were, me and another guy took her down by the bridge and did her.

Contrary to the State's posture, we find that the claim was sufficiently preserved for review. Defense counsel objected to Bullock's testimony on the grounds that the opinion of the witness was not relevant, that it was immaterial, and that it called for speculation. Defense counsel further stated that the witness "has not been qualified to interpret anything that he thought was said." The objection sufficiently apprised the court of the issue, namely, whether the witness could provide an opinion or inference as to what he thought Thorp meant. Thus, Thorp preserved this issue for appellate review.

As for the merits of Thorp's contention, we find that the trial court erred in allowing Bullock to testify as to what he believed Thorp's statement to mean. As a general rule, lay witnesses may not testify in the form of opinions or inferences; it is the function of the jury to draw those inferences. Cf. Kersey v. State, 73 Fla. 832, 840, 74 So. 983, 986 (1917); see also Charles W. Ehrhardt, Florida Evidence § 701.1, at 538 (1999 ed.). An exception to this rule is found in section 90.701, Florida Statutes, which permits a lay witness to proffer testimony in the form of an inference and opinion where:

(1) The witness cannot readily, and with equal accuracy and adequacy, communicate what he or she has perceived to the trier of fact without testifying in terms of inferences or opinions and the witness's use of inferences or opinions will not mislead the trier of fact to the prejudice of the objecting party; and

(2) The opinions and inferences do not require a special knowledge, skill, experience, or training.

§ 90.701, Fla. Stat. (1997); see also Kersey, 73 Fla. at 840, 74 So. at 986; Alexander v. State, 627 So.2d 35, 42 (Fla. 1st DCA 1993). Both prerequisites must be satisfied before a lay witness is allowed to testify in the form of an opinion or inference. See § 90.701; Ehrhardt,supra, at 540.

There is no claim in this case that Bullock's testimony required specialized skill or training. Accordingly, the second prong of section 90.701 is satisfied. However, it does not appear from the record that Bullock could not "readily, and with equal accuracy and adequacy, communicate what he . . . has perceived to the trier of fact without testifying in terms of inferences or opinions." Bullock had no difficulty conveying to the jury what Thorp said to him while in prison. He testified that Thorp admitted that he "did a hooker" and that he expected to be charged with murder. The exact meaning of Thorp's words and the inferences that could be drawn from them, however, were matters for the jury to consider. Accordingly, any inferences that may have been drawn from Thorp's statement that he "did a hooker" should have been made by the jury and not by the witness. See Kight v. State, 512 So.2d 922, 929 (Fla. 1987) (holding that there is no need to resort to lay opinion testimony where witness adequately conveyed his perception of defendant's words and actions to jury), receded from on other grounds, Davis v. State, 698 So.2d 1182 (Fla. 1997). Under these circumstances, there was no need to resort to testimony concerning Bullock's interpretation of Thorp's words. See id. Hence, the trial court erred in allowing this opinion testimony.

By permitting Bullock to interpret the meaning of Thorp's words, the trial court committed harmful error because it effectively turned Thorp's obvious admission of involvement in a crime into a confession of murder. As the record reflects, however, Thorp never confessed to murdering Sharon Case. Rather, during the penalty phase of the trial, he admitted only to having sexual intercourse with the victim, an act which is equally consistent with his admission to Bullock. Therefore, Bullock's opinion testimony that Thorp's statement that he "did a hooker" meant that he killed her, was undoubtedly prejudicial and we cannot say beyond a reasonable doubt that it did not affect the jury's verdict in this case. Under DiGuilio, the admission of Bullock's opinion testimony constitutes harmful error.

CONCLUSION

In sum, we find substantial error in the improper admission of certain highly prejudicial and damaging evidence against Thorp. We cannot find that error harmless under State v. DiGuilio, 491 So.2d 1129 (Fla. 1986). Accordingly, we reverse Thorp's conviction for first-degree murder and vacate his sentence of death. This case is remanded to the trial court with directions that a new trial be promptly conducted. Because we are remanding for a new trial, we need not address Thorp's remaining issues on appeal.

It is so ordered.

Shaw, Anstead, Pariente and Lewis, JJ., Concur.

Wells, C.J., Dissents with an opinion.

Quince, J., Dissents with an opinion, in which Harding, J., Concurs.